I made the decision to buy the expensive one based on the below bolded words. The cheaper one couldn't handle the size lens I plan on using. The 3.8" long 28mm-135mm IS lens. But I think I can fit the longer 70mm-300mm lens too. I'm hoping I can fit this into my book/computer bag I take to work. That way I can have it on me most of the time. I like the word modular. It's designed for change and improvement and the fact that one size does not fit all.
A case made from waterproof Cordura nylon fabric with dense closed-cell foam padding, made to carry a film or digital SLR camera.
• Pockets
* Large zippered front pocket will hold extra film and accessories
* Window Mesh pocket inside the top holds items like filters, memory cards, lens caps, and cleaning tissue.
• M.A.S. (Modular Accessory System) attachment slots allow you to add optional Tamrac accessories.
• S.A.S. (Strap Accessory System) has optional accessories that allow you to customize the bag
• Carrying options
* Adjustable, removable shoulder strap
* Padded carry handle
* Belt loop
• Compatible with compact digital or photo SLR camera with a lens attached (up to 5.5" long).